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Evidence of multiple gene interactions
Just as Wilhem Johannsen is conducting foundational work in plant genetics in Copenhagen, Swedish geneticist Herman Nilsson-Ehle is engaged in similar studies across the Oresund at the University of Lund. Nilsson-Ehle applies Mendelian principles to systematize the production of hybrid varieties of wheat. He finds that both qualitative traits (e.g., color) and quantitative characteristics (e.g., size) conform to Mendelian principles, but observes that the latter appear in quantitative gradients. This suggests that they result from interactions among multiple genes.